CEDAR RAPIDS, Iowa — An embattled northeast Iowa zoo is facing a second lawsuit – this time regarding treatment of its endangered African lions.

The Animal Legal Defense Fund and five Iowans filed a lawsuit Monday against the Cricket Hollow Zoo in Manchester, the Des Moines Register reported.

Earlier this year, the national group successfully sued the zoo to force the removal of endangered lemurs and tigers. The most recent lawsuit claims Cricket Hollow Zoo confines African lions in small, barren enclosures, disrupting and impairing the large cats.

Five tigers died at the zoo from 2013 through 2015, a California veterinarian testified in court last fall. The veterinarian said the tigers had received inadequate care and described the animals’ enclosures as covered in feces and old food.