WASHINGTON — Former White House national security adviser John Bolton said Monday that he is “prepared to testify” if he is subpoenaed by the Senate in its impeachment trial of President Trump, a surprise statement that bolstered Democrats in their push for new witnesses.

Bolton, who left the White House in September, said that he has weighed the issues of executive privilege and that after “careful consideration and study” decided that he would comply with any Senate subpoena.

“If the Senate issues a subpoena for my testimony, I am prepared to testify,” he said in a statement.

Senate Majority Leader Mitch McConnell has publicly expressed resistance to calling new witnesses in the upcoming trial, though Democrats are pressing to hear from Bolton and others who did not appear before the House in its impeachment inquiry.

House Speaker Nancy Pelosi has stalled the transmission of House-passed articles of impeachment against Trump in a bid for that witness testimony. Senate Democratic Leader Chuck Schumer has proposed calling several witnesses, including Bolton and Mulvaney, but McConnell has so far rejected Schumer’s terms.

It is uncertain when Pelosi will eventually send the articles to the Senate. If she decides to do so in the coming days, a trial could start as soon as this week.

In a tweet Monday, Pelosi did not address the timing. But she said Trump and McConnell have “run out of excuses” for not calling witnesses or producing documents Trump has blocked.

It’s unclear whether Bolton’s testimony would hurt or help the president. The two clashed while he was in the White House and offered differing versions of whether he resigned or was fired when he left office in September.

If Bolton were to appear, his testimony would give Congress and the public a highly anticipated, first-hand account from a Trump senior adviser who was present for key moments that have been described by others.

He’d almost certainly be asked, for instance, about a comment he was reported to have made to another White House adviser that he did not want to be “part of whatever drug deal” European Union ambassador Gordon Sondland and White House chief of staff Mick Mulvaney were “cooking up” as Trump pressured Ukraine to investigate Democrats.

That pressure, as Trump was withholding security aid to Ukraine, was at the heart of the inquiry in the House, which voted to impeach Trump on Dec. 18.

Schumer said in a statement after Bolton’s announcement that “momentum for uncovering the truth in a Senate trial continues.”

He said that it is “now up to four Senate Republicans to support bringing in Mr. Bolton” and the other witnesses he has proposed. Republicans hold the Senate with a 53-seat majority, and Democrats would have to find four Republicans to vote with them to support a subpoena.

Sen. Susan Collins, R-Maine, said it is too soon to decide what witnesses should be called, but that Bolton would be a possibility.

“There are a number of witnesses who may well be appropriate, of which John Bolton may certainly be one,” she said in a statement. “It is difficult to decide in isolation what the full universe of witnesses should look like before we have even heard the opening statements from the House managers and the President’s Counsel.”

Sen. Angus King, i-Maine, said in a recent op-ed that testimony from Bolton, as well as acting chief of staff Mick Mulvaney, Secretary of State Mike Pompeo and other White House officials, could go a long way toward clarifying whether Trump committed an impeachable offense.

“All could clear up, one way or another, the essential question of whether the president was interested in corruption generally in Ukraine last summer, or whether the withholding of military aid and the dangling of a White House meeting was motivated by his personal and political interest in publicized investigations of the 2016 election and the activities of one of his principal rivals,” King wrote for USA Today. “And this would be direct evidence, which no one can call hearsay.”

If any Republican opposes the subpoenas Democrats have proposed, Schumer said, “they would make absolutely clear they are participating in a cover up.”

McConnell has repeatedly called for a swift trial, with a possible final vote after opening arguments.

“We can’t hold a trial without the articles,” McConnell tweeted in the hours before Bolton’s statement. “The Senate’s own rules don’t provide for that. So, for now, we are content to continue the ordinary business of the Senate while House Democrats continue to flounder. For now.”

Bolton’s willingness to testify averts a potential legal standoff over whether close aides to the president can be forced to appear before Congress. Trump and his lawyers have claimed that those aides should not have to testify, arguing that they have special immunity, or executive privilege, not to.

The issue remains undecided in the courts now that a federal judge dismissed last week a lawsuit from Bolton’s former deputy, Charles Kupperman, who had asked the court whether he had to comply with a House subpoena or follow a White House orders that he not testify. U.S. District Judge Richard Leon dismissed the case as moot, noting that the House had withdrawn its subpoena for Kupperman and had said that it didn’t plan to reissue one.

The case had been closely watched for the repercussions it carried for Bolton’s testimony.

House Intelligence Committee Chairman Adam Schiff signaled in a tweet that he will not call Bolton again in the House, writing that “the Senate must allow testimony from him, Mulvaney and others.”

Schiff had invited Bolton to testify in the inquiry but had not subpoenaed him, saying he didn’t want the impeachment case to languish in courts. Instead, the House voted to impeach Trump for obstruction of Congress.

The Associated Press reported in November that Bolton is writing a book and has a deal with the publisher Simon & Schuster, according to three publishing officials with knowledge of the situation. Two said the deal was worth $2 million. They spoke on condition of anonymity because they weren’t authorized to speak publicly.

Trump tweeted Monday morning that the impeachment “hoax” must “end quickly.”

“It is a con game by the Dems to help with the Election!,” Trump tweeted.

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