KAMPALA, Uganda — South Sudan’s central government lost control of the capital of a key oil-producing state on Sunday, the military said, as renegade forces loyal to a former deputy president seized more territory in fighting that has raised fears of full-blown civil war in the world’s newest country.

Bentiu, the capital of oil-rich Unity state, is now controlled by a military commander loyal to former Vice President Riek Machar, said Col. Philip Aguer, the South Sudanese military spokesman.

“Bentiu is in the hands of a commander who has declared support for Machar,” he said. “Bentiu is not in our hands.”

The armed rebels were said to be in control days earlier of some of South Sudan’s oil fields, which have historically been a target for rebel movements, endangering the country’s economic lifeblood.

South Sudan gets nearly 99 percent of its government budget from oil revenues, and the country reportedly earned $1.3 billion in oil sales in just five months this year, according to the London-based watchdog group Global Witness.

Although the country’s capital, Juba, is mostly peaceful a week after a dispute among members of the presidential guard triggered violent clashes between military factions, fighting continues as the central government tries to assert authority in the states of Unity and Jonglei.

Bor, the capital of Jonglei, is said to be the scene of some of the fiercest clashes between government troops and rebels. Machar, now a fugitive wanted for his alleged role in what the government insists was an attempted coup last Sunday, has said in interviews that he is interested in negotiations that lead to the president’s exit. He denied involvement in a coup plot.

The U.N. Mission in South Sudan said in a statement Sunday that all non-critical staff in Juba are being evacuated to Uganda. The mission said the move was “a precautionary measure to reduce pressures on its limited resources” as it continues to provide assistance and shelter to over 20,000 civilians gathered inside its compounds in Juba, the mission said in a statement Sunday.

Hilde Johnson, the U.N. secretary-general’s envoy in South Sudan, said the evacuation does not mean the U.N. is “abandoning” South Sudan.

“We are here to stay, and will carry on in our collective resolve to work with and for the people of South Sudan,” she said. “To anyone who wants to threaten us, attack us or put obstacles in our way, our message remains loud and clear: we will not be intimidated.”

The United States and other countries have been evacuating their citizens from South Sudan, as violence escalates in the world’s newest country and threatens lives and oil production. Hundreds have been killed in the fighting and world leaders are concerned about full-blown civil war in a country with a history of ethnic violence and divided military loyalties.

On Saturday gunfire hit three U.S. military aircraft trying to evacuate American citizens in Bor, wounding four U.S. service members in the same region gunfire downed a U.N. helicopter on Friday.

Earlier this week the top military general in Bor defected with his troops, starting a rebellion that appears to be spreading to other parts of the country.

Aguer said Bor is still under the control of pro-Machar forces, disputing reports the rebels had fled as government troops advanced on Bor.

South Sudan’s President Salva Kiir, an ethnic Dinka, said on Monday that an attempted military coup had triggered the violence, and the blame was placed on Machar, an ethnic Nuer. But officials have since said a fight between Dinka and Nuer members of the presidential guard triggered the fighting that later spread across the East African country.

Machar’s ouster from the country’s No. 2 political position earlier this year had stoked ethnic tensions. Machar, who has criticized Kiir as a dictator, later said he would contest presidential elections in 2015.

U.N. Secretary-General Ban Ki-moon on Sunday urged South Sudan’s leaders “to do everything in their power” to stop the violence.

Foreign ministers from neighboring countries Kenya, Ethiopia, Uganda and Djibouti were in South Sudan earlier in the week to try and diffuse the crisis.

South Sudan, which became independent in 2011 after decades of a brutal war with Sudan, has been plagued by ethnic discord, corruption and conflict with Sudan over oil revenues.

Although the south inherited three-quarters of Sudan’s oil production when it declared independence in 2012, its oil exports are pumped through pipelines running north, raising concern a rebel takeover of southern oil fields could invite Sudan into the conflict.

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